Adrain, J.M., Westrop, S.R., Landing, E., Karim, T.S., Losso, S.R., Ng, R.Y., Bradley, A.B. & Pérez-Peris, F. (in press) Trilobite biostratigraphy of the upper Skullrockian Stage (Lower Ordovician; lower Tremadocian) on the northern Laurentian margin, western United States. Australasian Palaeontological Memoirs.
Adrain, J.M. (2025) The Early Ordovician bathyurid trilobites Licnocephala and Ibexocephala. Journal of Paleontology. Link (open access)
Pérez-Peris, F., Adrain, J.M. & Daley, A.C. (2024) Phylogenetics and systematics of the subfamilies Cheirurinae and Deiphoninae (Trilobita). Swiss Journal of Palaeontology, 143, 43. Link (open access)
Pérez-Peris, F., Adrain, J.M. & Allison C. Daley (2024) Ordovician paleobiogeography of the Suborder Cheirurina (Trilobita). Palaeogeography, Palaeoclimatology, Palaeoecology, 647, 112222. Link (open access)
Westrop, S.R., Welch, K.F., Engel, M.H. & Adrain, J.M. (2023) Faunal and paleoenvironmental changes at a Cambrian (Jiangshanian; Steptoean–Sunwaptan boundary interval) trilobite extinction event, in contrasting deep and shallow subtidal settings, Nevada and Oklahoma. Journal of Paleontology. Link (open access)
Adrain, J.M. & Pérez-Peris, F. (2023) Funeralaspis n. gen.: a new odontopleurine trilobite from the early Middle Ordovician (Dapingian) of Death Valley, eastern California, USA, and the classification of Ordovician odontopleurines. Zootaxa, 5336, 509–529.
Download PDF
Download PDF
Download PDF
Download PDF
Download PDF
Download PDF
Download PDF
Adrain, J.M., Karim, T.S. & Westrop, S.R. (2014) The Early Ordovician (Floian) bathyurid trilobite genera Jeffersonia, Cullisonia, and Bathyurina. Memoirs of the Association of Australasian Palaeontologists, 45, 305–347.
Download PDF
Download PDF
Download PDF
Westrop, S.R. & Adrain, J.M. (2013) Biogeographic shifts in a transgressive succession: The Cambrian (Furongian; Jiangshanian; latest Steptoean–earliest Sunwaptan) agnostoid arthropods Kormagnostella Romanenko and Biceragnostus Ergaliev in North America. Journal of Paleontology, 87, 804–817.
Adrain, J.M., McAdams, N.E.B. & Karim, T.S. (2012) The Middle Ordovician (Dapingian and Darriwilian) bathyurid trilobite Pseudoolenoides. Zootaxa, 3467, 1–97.
Adrain, J.M. & McAdams, N.E.B. (2012) The Lower Ordovician (upper Floian) bathyurid trilobite Aponileus Hu, with species from Utah, Texas, and Greenland. Zootaxa, 3293, 1–67.
Carlucci, J., Westrop, S.R., Amati, L., Adrain, J.M. & Swisher, R.E. (2012) A systematic revision of the Upper Ordovician trilobite genus Bumastoides (Trilobita: Illaenidae) with new species from Oklahoma, Virginia, and Missouri. Journal of Systematic Palaeontology, 10, 679–723.
Landing, E., Adrain, J.M., Westrop, S.R. & Kröger, B. (2012) Tribes Hill-Rochdale Formations in east Laurentia: proxies for Early Ordovician (Tremadoc) eustasy on a tropical passive margin (New York and west Vermont). Geological Magazine, 149, 93–123.
Adrain, J.M. (2011) Class Trilobita Walch, 1771. In: Zhang, Z.-Q. (ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 104–109.
McAdams, N.E.B. & Adrain, J.M. (2011) Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA. Zootaxa, 3144, 1-113.
Adrain, J.M., McAdams, N.E.B. & Westrop, S.R. (2011) Affinities of the Lower Ordovician (Tulean; lower Floian) trilobite Gladiatoria, with species from the Great Basin, western United States. Memoirs of the Association of Australasian Palaeontologists, 42, 321-367.
Adrain, J.M., McAdams, N.E.B., Westrop, S.R. & Karim, T. S. (2011) Systematics and affinity of the Lower Ordovician (Tulean; lower Floian) trilobite Psalikilopsis. Memoirs of the Association of Australasian Palaeontologists, 42, 369-416.
Adrain, J.M. (2011) A new trilobite biostratigraphy for the Lower Ordovician of western Laurentia and prospects for international correlation using pelagic trilobites. In: Gutiérrez-Marco, J. C., Rábano, I., Garcia-Bellido, D. (Eds.), Ordovician of the World. Cuadernos del Museo Geominero, 14, 41-42.
Landing, E., Westrop, S.R. & Adrain, J. M. (2011) The Lawsonian Stage - the Eoconodontus notchpeakensis (Miller, 1969) FAD and HERB carbon isotope excursion define a globally correlatable terminal Cambrian stage. Bulletin of Geosciences, 86, 621-640.
Westrop, S.R., Adrain, J.M. & Landing, E. (2011) The Cambrian (Sunwaptan, Furongian) agnostoid arthropod Lotagnostus Whitehouse, 1936, in Laurentian and Avalonian North America: systematics and biostratigraphic significance. Bulletin of Geosciences, 86, 569-594.
McAdams, N.E.B. & Adrain, J.M. (2011) Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA. Zootaxa, 2969, 1-68.
Adrain, T.S., Budd, A.F., & Adrain, J.M. (2011) Digitizing collections: Experiences from the University of Iowa Paleontology Repository digitization project. Geological Curator, 9, 291-299.
McAdams, N.E.B., & Adrain, J.M. (2011) Systematics of the Lower Ordovician pliomerid trilobite Hintzeia, with species from the Great Basin, western USA. Zootaxa, 2910, 1-45.
Westrop, S.R., Waskiewicz, R.A., & Adrain, J.M. (2010) Systematics of Dokimocephalus and related trilobites from the Late Cambrian (Steptoean; Millardan and Furongian series) of Laurentian North America. Journal of Systematic Palaeontology, 8, 545-606.
McAdams, N.E.B., & Adrain, J. M. (2010) A new species of the Lower Ordovician pliomerid trilobite Pseudocybele and its biostratigraphic significance. Zootaxa, 2550, 21-38.
Adrain, J.M., McAdams, N.E.B. & Westrop, S.R. (2009) Trilobite biostratigraphy and revised bases of the Tulean and Blackhillsian Stages of the Ibexian Series, Lower Ordovician, western United States. Memoirs of the Association of Australasian Palaeontologists, 37, 5435-58.
McAdams, N.E.B. & Adrain, J.M. (2009) Heckethornia, a new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA. Canadian Journal of Earth Sciences, 46, 875-914.
McAdams, N.E.B. & Adrain, J.M. (2009) New pliomerid trilobite genus Lemureops from the Lower Ordovician (Ibexian; Tulean, Blackhillsian) of western Utah, USA. Memoirs of the Association of Australasian Palaeontologists, 37, 491-540.
Westrop, S.R. & Adrain, J.M. (2009) The late Cambrian (Furongian; Steptoean) trilobite genus Xenocheilos Wilson, 1949: systematics and biostratigraphic significance. Memoirs of the Association of Australasian Palaeontologists, 37, 351-368.
Adrain, J.M., Peters, S.E. & Westrop, S.R. (2009) The Marjuman trilobite Cedarina Lochman: thoracic morphology, systematics, and new species from western Utah and eastern Nevada, USA. Zootaxa, 2218, 35-58.
Adrain, J.M. (2009) New and revised species of the aulacopleurid trilobite Maurotarion from the Lower Devonian (Pragian) of Nevada. Zootaxa, 2215, 1-23.
Westrop, S.R. & Adrain, J.M. (2009) The Late Cambrian (Steptoean; Furongian) trilobite Pseudokingstonia Palmer, 1965 in North America. Canadian Journal of Earth Sciences, 46, 355-360.
Adrain, J.M., Chatterton, B.D.E. & Kloc, G.J. (2008) Systematics of the koneprusiine trilobites, with new taxa from the Silurian and Devonian of Laurentia. Journal of Paleontology, 82, 657-675.
Westrop, S.R., Eoff, J.D., Ng, T.-W., Dengler, A.A. & Adrain, J.M. (2008) Classification of the Late Cambrian (Steptoean) trilobite genera Cheilocephalus Berkey, 1898 and Oligometopus Resser, 1936 from Laurentia. Canadian Journal of Earth Sciences, 45, 725-744.
Adrain, J.M. (2008) A global species database of Trilobita: progress, results, and revision of the Treatise. In: Rábano, I., Gozalo, R. & Garcia-Bellido, D. (Eds.), Advances in trilobite research. Cuadernos del Museo Geominero, 9, 27-28.
McAdams, N.E.B. & Adrain, J.M. (2008) A new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian) of western Utah, USA. In: Rábano, I. & Garcia-Bellido, D. (Eds.), A new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian) of western Utah, USA. Cuadernos del Museo Geominero, 9, 251-252.
Adrain, J.M. & Westrop, S.R. (2007) Bearriverops, a new Lower Ordovician trilobite genus from the Great Basin, western USA, and classification of the family Dimeropygidae. Canadian Journal of Earth Sciences, 44, 337-366.
Adrain, J.M. & Westrop, S.R. (2007) The hystricurid trilobite Metabowmania in the Lower Ordovician (Ibexian; Stairsian) of the Great Basin, Utah and Idaho, USA. Memoirs of the Association of Australasian Palaeontologists, 34, 227-242.
Westrop, S.R. & Adrain, J.M. (2007) Bartonaspis new genus, a trilobite species complex from the base of the Upper Cambrian Sunwaptan Stage in North America. Canadian Journal of Earth Sciences, 44, 987-1003.
Westrop, S.R., Waskiewicz, R.A. & Adrain, J.M. (2007) The Late Cambrian (Steptoean) trilobite genus Bynumina Resser, 1942, in North America. Memoirs of the Association of Australasian Palaeontologists, 34, 357-376.
Adrain, J.M. & Tetreault, D.K. (2006) The brachymetopid trilobite Radnoria in the Silurian (Wenlock) of New York State and Arctic Canada. Canadian Journal of Earth Sciences, 42, 2087-2096. (for 2005)
Adrain, J.M. & Westrop, S.R. (2006) New genus of dimeropygid trilobites from the earliest Ordovician of Laurentia. Acta Palaeontologica Polonica, 51, 541-550.
Adrain, J.M. & Westrop, S.R. (2006) Notchpeakia, a new genus of Upper Cambrian (Sunwaptan) "entomaspidid" trilobites. Journal of Paleontology, 80, 1152-1171.
Adrain, J.M. & Westrop, S.R. (2006) New earliest Ordovician trilobite genus Millardicurus: The oldest known hystricurid. Journal of Paleontology, 80, 650-671.
Adrain, J.M. (2005) Aulacopleurid trilobites from the Upper Ordovician of Virginia. Journal of Paleontology, 79, 542-563.
Adrain, J.M. & Westrop, S.R. (2005) Lower Ordovician trilobites from the Baumann Fiord Formation, Ellesmere Island, Arctic Canada. Canadian Journal of Earth Sciences, 42, 1523-1546.
Adrain, J.M. & Westrop, S.R. (2005) Late Cambrian ptychaspidid trilobites from western Utah: implications for trilobite systematics and biostratigraphy. Geological Magazine, 142, 377-398.
Adrain, J.M. & Westrop, S.R. (2004) A Late Cambrian (Sunwaptan) silicified trilobite fauna from Nevada. Bulletins of American Paleontology, 365, 1-51.
Adrain, J.M., Edgecombe, G.D., Fortey, R.A., Laurie, J., McCormick, T., Owen, A.W., Waisfeld, B.G., Webby, B.D., Westrop, S.R. & Zhou, Z.-Y. (2004) Trilobites. In: Webby, B.D., Droser, M.L. & Paris, F. (Eds.), The Great Ordovician Biodiversification Event. Columbia University Press, New York, pp. 231-254.
Adrain, J.M., Lee, D.-C., Westrop, S.R., Chatterton, B.D.E. & Landing, E. (2003) Classification of the trilobite subfamilies Hystricurinae and Hintzecurinae subfam. nov., with new genera from the Lower Ordovician (Ibexian) of Idaho and Utah. Memoirs of the Queensland Museum, 48, 553-586.
Jell, P.A. & Adrain, J.M. (2003) Available generic names for trilobites. Memoirs of the Queensland Museum, 48, 331-553.
Download PDF
Adrain, J.M. & Westrop, S.R. (2003) Paleobiodiversity: We need new data. Paleobiology, 29, 22-25.
Download PDF
Adrain, J.M. (2003) Validity and composition of the Silurian trilobite genera Borealarges and Dicranogmus, with new species from the Canadian Arctic. Canadian Journal of Earth Sciences, 40, 749-763.
Download PDF
Adrain, J.M., Westrop, S.R., Landing, E. & Fortey, R.A. (2001) Systematics of the Ordovician trilobites Ischyrotoma and Dimeropygiella, with species from the type Ibexian area, western U.S.A. Journal of Paleontology, 75, 947-971.
Download PDF
Adrain, J.M. & Westrop, S.R. (2001) Stratigraphy, phylogeny, and species sampling in time and space. In: Adrain, J.M., Edgecombe, G.D. & Lieberman, B.S. (Eds.), Fossils, Phylogeny, and Form: An Analytical Approach. Kluwer, New York, pp. 291-322.
Download PDF
Adrain, J.M. (2001) Systematic paleontology. Journal of Paleontology, 75, 1055-1057.
Download PDF
Download PDF
Brochu, C.A., Bryant, H.N., Theodor, J.M., O'Leary, M.A., Adrain, J.M. & Sumrall, C.D. (2001) Modern phylogenetics in paleontology: comments on Vermeij, 1999. Paleobiology, 27, 174-176.
Download PDF
Adrain, J.M. & Westrop, S.R. (2000) An empirical assessment of taxic paleobiology. Science, 289, 110-112.
Download PDF
Adrain, J.M., Westrop, S.R., Chatterton, B.D.E. & Ramsköld, L. (2000) Silurian trilobite alpha diversity and the end-Ordovician mass extinction. Paleobiology, 26, 625-646.
Download PDF
Adrain, J.M. & Westrop, S.R. (1999) Trilobite paleobiology: Past, present, and future. Journal of Paleontology, 73, 161-163.
Download PDF
Hughes, N.C., Chapman, R.E. & Adrain, J.M. (1999) The stability of thoracic segmentation in trilobites: a case study in developmental and ecological constraints. Evolution and Development, 1, 24-35.
Download PDF
Adrain, J.M., Fortey, R.A. & Westrop, S.R. (1998) Post-Cambrian trilobite diversity and evolutionary faunas. Science, 280, 1922-1925.
Download PDF
Download PDF
Westrop, S.R. & Adrain, J.M. (1998) Trilobite diversity and the Ordovician Radiation: a reply to Miller et al. Paleobiology, 24, 529-533.
Download PDF
Westrop, S.R. & Adrain, J.M. (1998) Trilobite alpha diversity and the reorganization of Ordovician benthic marine communities. Paleobiology, 24, 1-16.
Download PDF
Davidek, K., Landing, E., Bowring, S.A., Westrop, S.R., Rushton, A.W.A., Fortey, R.A. & Adrain, J.M. (1998) New uppermost Cambrian U-Pb date from Avalonian Wales and age of the Cambrian-Ordovician boundary. Geological Magazine, 135, 305-309.
Download PDF
Adrain, J.M. (1997) Proetid trilobites from the Silurian (Wenlock-Ludlow) of the Cape Phillips Formation, Canadian Arctic Archipelago. Palaeontographia Italica, 84, 21-111.
Download PDF
(extremely homemade, but better than nothing as original is difficult to find)
Adrain, J.M. & Fortey, R.A. (1997) Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the Natural History Museum, London, Geology Series, 53, 79-115.
Download PDF
(homemade)
Adrain, J.M. & Ramsköld, L. (1997) Silurian Odontopleurinae (Trilobita) from the Cape Phillips Formation, Arctic Canada. Journal of Paleontology, 71, 237-261.
Download PDF
Adrain, J.M. & Edgecombe, G.D. (1997) Silurian (Wenlock) calymenid trilobites from the Cape Phillips Formation, central Canadian Arctic. Journal of Paleontology, 71, 657-682.
Download PDF
Adrain, J.M. & Edgecombe, G.D. (1997) Silurian encrinurine trilobites from the central Canadian Arctic. Palaeontographica Canadiana, 14, 1-109.
Download PDF
(homemade)
Adrain, J.M. & Kloc, G.J. (1997) Lower Devonian aulacopleuroidean trilobites from Oklahoma. Journal of Paleontology, 71, 703-712.
Download PDF
Adrain, J.M. (1996) A new otarionine trilobite from the Henryhouse Formation (Silurian, Ludlow) of Oklahoma. Journal of Paleontology, 70, 611-614.
Download PDF
Adrain, J.M. & Ramsköld, L. (1996) The lichid trilobite Radiolichas in the Silurian of Arctic Canada and Gotland, Sweden. Geological Magazine, 133, 147-158.
Download PDF
Adrain, J.M. & MacDonald, E.W. (1996) Phacopid trilobites from the Silurian of Arctic Canada. Journal of Paleontology, 70, 1091-1094.
Download PDF
Adrain, J.M. & Chatterton, B.D.E. (1996) The otarionine trilobite Cyphaspis, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 70, 100-110.
Download PDF
Adrain, J.M. & Edgecombe, G.D. (1996) Devonian aulacopleurid trilobites of the Malvinokaffric Realm. Geobios, 29, 417-436.
Download PDF
Adrain, J.M., Chatterton, B.D.E. & Blodgett, R.B. (1995) Silurian trilobites from southwestern Alaska. Journal of Paleontology, 69, 723-736.
Download PDF
Adrain, J.M. & Edgecombe, G.D. (1995) Balizoma and the new genera Aegrotocatellus and Perirehaedulus: encrinurid trilobites from the Douro Formation (Silurian, Ludlow) of the central Canadian Arctic. Journal of Paleontology, 69, 736-752.
Download PDF
Adrain, J.M. & Chatterton, B.D.E. (1995) The otarionine trilobites Harpidella and Maurotarion, with species from northwestern Canada, the United States, and Australia. Journal of Paleontology, 69, 307-326.
Download PDF
Adrain, J.M. & Chatterton, B.D.E. (1995) Aulacopleurine trilobites from the Llandovery of northwestern Canada. Journal of Paleontology, 69, 326-340.
Download PDF
Edgecombe, G.D. & Adrain, J.M. (1995) Silurian calymenid trilobites from the United States. Palaeontographica Abteilung A, 235, 1-19.
No PDF available (sorry!)
Adrain, J.M. (1994) The lichid trilobite Borealarges n. gen., with species from the Silurian of Arctic Canada. Journal of Paleontology, 68, 1081-1099.
Download PDF
Adrain, J.M. & Chatterton, B.D.E. (1994) The aulacopleurid trilobite Otarion, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 68, 305-323.
Download PDF
Adrain, J.M. & Wilson, M.V.H. (1994) Early Devonian cephalaspids (Vertebrata: Osteostraci: Cornuata) from the southern Mackenzie Mountains, N.W.T., Canada. Journal of Vertebrate Paleontology, 14, 301-319.
Download PDF
Ramsköld, L., Adrain, J.M., Edgecombe, G.D. & Siveter, D.J. (1994) Silurian calymenid trilobite Alcymene n. gen., with new species from the Ludlow of Gotland, Sweden. Journal of Paleontology, 68, 556-569.
Download PDF
Adrain, J.M. & Chatterton, B.D.E. (1993) A new rorringtoniid trilobite from the Ludlow of Arctic Canada. Canadian Journal of Earth Sciences, 30, 1634-1643.
Download PDF
(homemade)
Adrain, J.M. (1992) Machaeridian classification. Alcheringa, 16, 15-32.
Download PDF
(homemade)
Adrain, J.M., Chatterton, B.D.E. & Cocks, L.R.M. (1991) A new species of machaeridian from the Silurian of Podolia, USSR, with a review of the Turrilepadidae. Palaeontology, 34, 637-651.
Download PDF
Adrain, J.M. & Chatterton, B.D.E. (1990) Odontopleura (Trilobita, Silurian), and a method of constrained congruency analysis. Journal of Paleontology, 64, 600-614.
Download PDF